Stoneflies belong to the order Plecoptera ('downwings'). Like the mayflies, these aquatic insects have been around for over 200 million years - 100 million years before the earliest recognizable ancestors of salmon and trout appeared on earth.

Stonefly nymphs are common inhabitants of the fast-flowing rocky stretches of streams (hence the name) and are usually restricted to well oxygenated, cold, tumbling mountain streams or riffle sections of cold unpolluted waters. A few species are also found in northern or cold lakes which have considerable wave action.

Key features of stonefly nymphs include:
- a flattened to cylindrical slender body
- distinct wing pads
- well developed and widely separated eyes
- long slender antennae
- two tails
- six well developed legs

Depending on species and habitat, stonefly nymphs come in various shades of yellow, orange, brown and black.

Stoneflies have what is known as an incomplete lifecycle which consists of three stages; namely, egg-nymph (larvae)-adult. Nymphal development may be one year, occasionally greater.

When mature, the nymph crawls from the stream onto land where it molts to emerge as a winged adult. The adults resemble the nymphs, but in addition have two pairs of elongated, heavily veined, wings which typically are held flat or occasionally wrapped partially around the abdomen at rest. A few species have reduced or absent wings.

Adults tend to be weak fliers, which presumably explains why they spend so much time hiding under logs or rocks, or clinging under leaves and other streamside vegetation. Adult stoneflies tend to live longer than adult mayflies; but not by much. Instead of days, most adult stoneflies live a few weeks. During this short time, the adults mate, and the females lay eggs. Some species lay eggs by dive bombing into fast flowing waters; others simply crawl back underwater to deposit thier eggs.

Stoneflies are generally recognized as being important food source for trout and salmon inhabiting coastal streams - they are present year round, and they can be found in the highest alpine streams down to the estuary. The flyfisher can expect to find at least 60 kinds (species) of stoneflies in Western coastal streams. It is not necessary to learn to recognize all the stonefly species; nor is it that useful to memorize the various generic groups, but I do think it is worth while to have a simple working classification for fishing purposes. One such classification, developed for species found in the Bella Coola river is listed below, together with emergence times.

1. Little Green and Yellow Stoneflies:
Alloperla species April-September
Kathroperla perdita May-July
Naeviperla forcipata July-September
Paraperla frontalis April-August
Suwallia pallidula May-October
Sweltsa spp May-September
Triznaka diversa April-September
Size: 10-17 mm
Colour of pattern: Yellow or yellow-green; few brownish.

2. Little Black Stoneflies:
Capnia species February-April
Eucapnopsis brevicauda February-July
Isocapnia species February-April
Size: most 3-6 mm; few species to 20 mm
Colour of pattern: Black to dark reddish brown.

3. Big Golden Stoneflies:
Classenia sabulosa May-June
Hesperoperla pacifica May-June
Size: 25-38 mm
Colour of pattern: golden-yellow to yellowish-brown.

4. Rolled-winged Stoneflies:
Despaxia augusta Summer-Fall
Leuctra spp February-April
Paraleuctra spp March-August
Size: 9-13 mm
Colour of pattern: dark brown to black.

5. Medium Brown and Yellow Stoneflies:
Diura knowltoni April-June
Isogenoides colubrinus March-August
Isoperla spp April-October
Kogotus nonus April-September
Megarcys spp April-August
Rickera sorpta Spring-Summer
Skwala parallela Spring-Summer
Size: 8-25 mm
Colour of pattern: brown with yellow stripes or bands.

6. Early Black and Brown Stoneflies:
Doddsia occidentalis Winter-Spring
Toenionema spp Winter-Spring
Size: 7-13 mm
Colour of pattern: dark brown to black.

7. Little Brown Stoneflies:
Malenka spp Spring-Summer
Podmosta spp Spring-Summer
Prestoia besametsa Spring-Summer
Visola cataractae Spring-Summer
Zapada spp February-August
Size: 3-10 mm
Colour of pattern: brown to black.

8. Giant Stonefly (Salmonfly):
Pteronarcys californicus April-August
Size: 25-50 mm
Colour of pattern: Dark brown with orangy underbody.

This simple classification is particularly useful because all one has to do is recognize that the insect is a stonefly, then the pattern used is based on color of the nymph or adult.

By Harvey Thommasen with photography by Mike Wigle